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Silo was a world-famous male chinstrap penguin in New York City’s Central Park Zoo, who, along with his male partner Roy, became known as the “gay penguins” after being featured in a New York Times article in 2004.

Hatched in 1987, Silo met Roy at the Central Park Zoo in 1998, and the two took to each other almost immediately. Zookeepers and visitors alike noticed how smitten the birds were, as they engaged in mating rituals with one another such as entwining their necks and squawking mating calls. The following year, the couple attempted to hatch a rock as though it were an egg, which inspired zookeepers to give them first a dummy egg, and then an actual penguin egg that another couple was unable to hatch.

Silo and Roy successfully incubated the egg and hatched a female chick that was named Tango, who went on to pair with another female penguin called Tanuzi. The story captured the hearts of millions across the world.

Not long after their initial media coverage propelled them to fame, Silo and Roy were ousted from their nest by a more aggressive pair of penguins in May 2004 and subsequently began to drift apart. It would seem like their story was at an end, but, in true dramatic fashion, there was a plot twist. And, as it turned out, the penguins weren’t both gay.

Roy found another male partner named Blue. Silo, however, ended up bonding with a female penguin named Scrappy, demonstrating that he was, in fact, bisexual. Given how widespread bisexuality is in nature, and how rare exclusive homosexuality is among non-human animals, it shouldn’t surprise us that one of the “gay penguins” was actually a bi penguin.

Silo and Roy’s story made an outsized splash in American culture. It inspired the children’s book And Tango Makes Three (2005), which was a Lambda Literary Award finalist that year. According to the American Library Association, And Tango Makes Three became one of the most banned children’s books in the United States over its depiction of a same-sex relationship. Nevertheless, it became a bestseller. Silo and Roy have also been featured in several theatrical productions, such as the 2011 play Birds of a Feather, and a 2011 play/ballet version of And Then Came Tango.

The whole saga also got wrapped up in the American culture wars of the era. It ruffled the feathers of some Christian conservatives and also emboldened many in the LGBT community to criticize the attempts of other zoos to forcibly separate same-sex animal couples.

Roy and Tango have both died in the years since their story broke in 2004. Astonishingly, as of 2018, Silo was still reportedly alive and had fathered a biological chick of his own! His current status is unknown, but given that the life expectancy of chinstrap penguins is 20–22 years for those in captivity, Silo has likely passed away as a very old bisexual penguin.